St John's College Oxford
Professor Alan Grafen

Professor Alan Grafen FRS

Tutorial Fellow in Quantitative Biology Senior Tutor



I teach population biology in the first year, and statistics and behavioural ecology in the second and third years of the Biological Sciences course.  I also teach Animal Behaviour to Human Scientists at St John's.

Research Interests

My research involves applying mathematical and logical theorising to evolutionary problems. I have previously worked on animal signals, sexual selection, kin selection, kin recognition, the statistics of cross-species comparisons and optimality models of sequences of behaviour. My current research program is my 'formal Darwinism project', which aims to capture Darwin's central argument about evolution by natural selection in a mathematical framework. With a dozen papers published since 1999, this program is about half-way through.